Central and box coolers take the seawater side, and the seawater side grows things
Plate central coolers on a seawater duty foul biologically as well as chemically. Larvae settle in the ports and in the low-velocity lanes of the plate pack, then grow, and the pressure drop climbs long before the approach temperature tells you anything is wrong. Box coolers in the sea chest are worse again, because the growth is on the outside of the tube nest where nothing circulates. Titanium plates tolerate the chemistry that shifts it; the gaskets and the frame studs often do not, which is what limits how aggressive a clean can be.
